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Disclosery

RYN RAYONIER INC

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REIT (common), issuer CIK 52827

327 managers (336 13F filers) reported a position in RYN shares at 2025-06-30, 145,272,194 shares in all. 43 filers opened a position and 66 filers sold out during the quarter.

Managers
327
13F filers
336
New
43
Added
135
Reduced
96
Sold out
66
Shares held by filers
145,272,194

Shares held by filers adds up every 13F filer's report, so a share reported by a parent and by its subsidiary counts twice (H9). Managers counts each filer group once.

13D and 13G filers

Each filer's latest stake. The shares are the aggregate the reporting persons state, which can include entities that file no 13F.

Insider trades

The latest Form 4 transactions, non-derivative.
OwnerDateSharesPriceFiling
Daniels Ryan M.officer (SVP, Wood Products)2026-10-02-14 S$18.22Form 4

13F holders, Q2 2025

Filers in the available 13F reports with a share position at 2025-06-30, filer groups collapsed to one row (method), largest first. Options and principal amounts are left out. A group row states no % of portfolio, since each member reports its own portfolio: the members' percents are listed under it. A value about 1,000 times or 1/1,000 of the price other filers imply is shown as filed but neither ranks the row nor enters a sum.

HolderSharesValue% of portfolioChangeSource
VANGUARD GROUP INCgroup of 2 filers holding20,975,711$465M+88,476Timeline
PRICE T ROWE ASSOCIATES INC /MD/16,909,037$375M0.04%-71,11713F-HRTimeline
BlackRock, Inc.in group BlackRock, Inc.14,760,570$327M0.01%-1,316,28813F-HRTimelineGroup
NORGES BANK13,730,279$305M0.04%013F-HRTimeline
COHEN & STEERS, INC.in group COHEN & STEERS, INC.13,035,239$289M0.53%+571,69613F-HRTimelineGroup
BRITISH COLUMBIA INVESTMENT MANAGEMENT …4,842,552$107M0.62%+639,00013F-HRTimeline
STATE STREET CORPin group STATE STREET CORP4,566,148$101M<0.01%-123,79413F-HRTimelineGroup
Legal & General Group Plcin group Legal & General Group Plc4,193,530$93M0.02%+823,31113F-HRTimelineGroup
Pictet Asset Management Holding SAin group Pictet Asset Management Holdi…3,889,913$86.3M0.09%-130,24313F-HRTimelineGroup
JPMORGAN CHASE & COin group JPMORGAN CHASE & CO3,688,309$81.8M0.01%+754,07613F-HRTimelineGroup
VICTORY CAPITAL MANAGEMENT INC2,962,019$65.7M0.04%+446,72113F-HRTimeline
GEODE CAPITAL MANAGEMENT, LLCin group GEODE CAPITAL MANAGEMENT, LLC2,833,226$62.8M<0.01%-4,26813F-HRTimelineGroup
CHARLES SCHWAB INVESTMENT MANAGEMENT INCin group Charles Schwab Trust Co2,646,007$58.7M0.01%+38,79613F-HRTimelineGroup
Impax Asset Management Group plcin group Impax Asset Management Group …2,484,057$54.7M0.31%-2,743,78713F-HRTimelineGroup
AQR CAPITAL MANAGEMENT LLCin group AQR CAPITAL MANAGEMENT LLC1,785,840$39.3M0.03%+279,72113F-HRTimelineGroup
Global Alpha Capital Management Ltd.1,700,299$37.7M2.35%+308,20013F-HRTimeline
NORTHERN TRUST CORPin group NORTHERN TRUST CORP1,678,329$37.2M0.01%+96,10213F-HRTimelineGroup
FRANKLIN RESOURCES INCin group Franklin Resources1,423,859$31.6M0.01%-81,93513F-HRTimelineGroup
Central Securities Corp1,201,474$26.6M2.35%013F-HRTimeline
UBS Group AGgroup of 2 filers holding1,137,378$25.2M+58,104Timeline
FULLER & THALER ASSET MANAGEMENT, INC.1,048,051$23.2M0.08%+5,82013F-HRTimeline
Bank of New York Mellon Corpin group Bank of New York Mellon Corp1,040,917$23.1M<0.01%-93,04913F-HRTimelineGroup
Universal- Beteiligungs- und Serviceges…in group Universal- Beteiligungs- und …931,871$20.7M0.04%+86,03813F-HRTimelineGroup
Invesco Ltd.in group Invesco Ltd.863,707$19.2M<0.01%+124,57413F-HRTimelineGroup
Nuveen, LLCin group TIAA822,868$18.3M0.01%-116,41913F-HRTimelineGroup
Qube Research & Technologies Ltd790,239$17.5M0.02%+723,73213F-HRTimeline
MILLENNIUM MANAGEMENT LLCin group MILLENNIUM MANAGEMENT LLC663,626$14.7M0.01%-842,88013F-HRTimelineGroup
ARROWSTREET CAPITAL, LIMITED PARTNERSHIPin group ARROWSTREET CAPITAL, LIMITED …657,158$14.6M0.01%-30,18413F-HRTimelineGroup
VOLORIDGE INVESTMENT MANAGEMENT, LLC645,477$14.3M0.05%+469,53313F-HRTimeline
FIRST TRUST ADVISORS LPin group FIRST TRUST ADVISORS LP582,595$12.9M0.01%+184,75413F-HRTimelineGroup
MORGAN STANLEYin group MORGAN STANLEY572,579$12.7M<0.01%-313,78113F-HRTimelineGroup
CITADEL ADVISORS LLCin group CITADEL ADVISORS LLC520,395$11.5M0.01%+55913F-HRTimelineGroup
Balyasny Asset Management L.P.in group Balyasny Asset Management505,351$11.2M0.03%+457,35213F-HRTimelineGroup
Russell Investments Group, Ltd.in group Russell Investments Group, Lt…505,240$11.2M0.01%-68,06513F-HRTimelineGroup
Ancora Advisors LLCin group Ancora Advisors LLC472,278$10.5M0.22%+99,38413F-HRTimelineGroup
Focus Partners Wealthgroup of 2 filers holding451,284$10M+14,469Timeline
Invenomic Capital Management LP446,135$9.9M0.56%new13F-HRTimeline
GOLDMAN SACHS GROUP INCin group GOLDMAN SACHS GROUP INC433,515$9.62M<0.01%+58,68113F-HRTimelineGroup
RHUMBLINE ADVISERS433,159$9.61M0.01%-4,26413F-HRTimeline
MANUFACTURERS LIFE INSURANCE COMPANY, T…in group MANUFACTURERS LIFE INSURANCE …385,085$8.54M0.01%+63,03213F-HRTimelineGroup
Point72 Asset Management, L.P.in group Point72 Asset Management, L.P.372,000$8.25M0.02%+95,65013F-HRTimelineGroup
BANK OF AMERICA CORP /DE/in group BANK OF AMERICA CORP /DE/352,412$7.82M<0.01%-94,73113F-HRTimelineGroup
Crossmark Global Holdings, Inc.331,283$7.35M0.12%+1,49613F-HRTimeline
California Public Employees Retirement …321,094$7.12M<0.01%+19,98613F-HRTimeline
PRINCIPAL FINANCIAL GROUP INCgroup of 2 filers holding318,631$7.07M-5,394Timeline
ALLIANCEBERNSTEIN L.P.311,626$6.91M<0.01%+63,23413F-HRTimeline
Swiss National Bank283,376$6.29M<0.01%+9,20013F-HRTimeline
Gotham Asset Management, LLC277,230$6.15M0.04%+201,59113F-HRTimeline
THIRD AVENUE MANAGEMENT LLC258,129$5.73M1.18%-37013F-HRTimeline
FIRETRAIL INVESTMENTS PTY LTD239,578$5.29M1.38%-177,62613F-HRTimeline
IEQ CAPITAL, LLC230,487$5.11M0.03%+71,86113F-HRTimeline
Robeco Institutional Asset Management B…in group Robeco Institutional Asset Ma…228,050$5.06M0.01%+16,00013F-HRTimelineGroup
NORDEA INVESTMENT MANAGEMENT ABin group NORDEA INVESTMENT MANAGEMENT …225,389$4.96M<0.01%+33,07113F-HRTimelineGroup
RENAISSANCE TECHNOLOGIES LLC222,450$4.93M0.01%+62,60013F-HRTimeline
D. E. Shaw & Co., Inc.in group D. E. Shaw & Co., Inc.190,384$4.22M<0.01%-103,93913F-HRTimelineGroup
TWO SIGMA INVESTMENTS, LPgroup of 2 filers holding188,030$4.17M-300,741Timeline
Teramo Advisors, LLC175,720$3.9M1.42%013F-HRTimeline
Retirement Systems of Alabama171,355$3.8M0.01%+32113F-HRTimeline
NEW YORK STATE TEACHERS RETIREMENT SYST…169,711$3.76M0.01%+3,59213F-HRTimeline
STATE BOARD OF ADMINISTRATION OF FLORID…160,400$3.56M0.01%+1,14013F-HRTimeline
PRUDENTIAL FINANCIAL INCin group PRUDENTIAL FINANCIAL INC156,988$3.48M<0.01%+79,55313F-HRTimelineGroup
Holocene Advisors, LP155,928$3.46M0.01%+23,83513F-HRTimeline
ROYAL BANK OF CANADAin group ROYAL BANK OF CANADA151,249$3.35M<0.01%-7,86713F-HR/ATimelineGroup
BARCLAYS PLCin group BARCLAYS PLC149,283$3.31M<0.01%-57,81713F-HR/ATimelineGroup
BANK OF MONTREAL /CAN/in group BANK OF MONTREAL /CAN/149,083$3.31M<0.01%+127,70513F-HRTimelineGroup
Verition Fund Management LLC144,963$3.22M0.02%+93,42813F-HRTimeline
WELLS FARGO & COMPANY/MNin group WELLS FARGO & COMPANY/MN138,905$3.08M<0.01%+108,13513F-HRTimelineGroup
AMERIPRISE FINANCIAL INCin group AMERIPRISE FINANCIAL INC136,661$3.03M<0.01%-7,57613F-HRTimelineGroup
BNP PARIBAS FINANCIAL MARKETSin group BNP PARIBAS FINANCIAL MARKETS130,716$2.9M<0.01%+90,34713F-HRTimelineGroup
CALIFORNIA STATE TEACHERS RETIREMENT SY…127,688$2.83M<0.01%013F-HRTimeline
Uniplan Investment Counsel, Inc.116,813$2.59M0.26%-2,27913F-HRTimeline
Caisse de depot et placement du Quebec106,492$2.36M0.01%new13F-HRTimeline
ALGERT GLOBAL LLC99,370$2.2M0.05%+27,56013F-HR/ATimeline
RAYMOND JAMES FINANCIAL INCin group RAYMOND JAMES FINANCIAL INC94,830$2.1M<0.01%+33,44113F-HRTimelineGroup
State of Alaska, Department of Revenue91,510$2.03M0.02%-25313F-HRTimeline
Cerity Partners LLCin group Cerity Partners LLC90,548$2.01M<0.01%+22,02013F-HRTimelineGroup
NEW YORK STATE COMMON RETIREMENT FUND88,862$1.97M<0.01%013F-HR/ATimeline
Sequoia Financial Advisors, LLC88,519$1.96M0.01%+6,21713F-HRTimeline
Tidal Investments LLCin group Tidal Investments LLC88,462$1.96M0.01%-9,66213F-HRTimelineGroup
CANADA LIFE ASSURANCE Coin group CANADA LIFE ASSURANCE Co85,751$1.9M<0.01%-3,78113F-HRTimelineGroup
State of New Jersey Common Pension Fund…in group State of New Jersey Common Pe…80,696$1.79M0.01%+6,54813F-HRTimelineGroup
HighTower Advisors, LLC80,315$1.78M<0.01%+1,62313F-HRTimeline
State of Tennessee, Department of Treas…78,910$1.75M0.01%013F-HRTimeline
MetLife Investment Management, LLCin group MetLife Investment Management…75,738$1.68M0.01%-2,53913F-HRTimelineGroup
Cubist Systematic Strategies, LLC75,572$1.68M0.02%+47,12613F-HRTimeline
Treasurer of the State of North Carolina72,320$1.6M0.01%+1,78313F-HRTimeline
BFSG, LLC72,271$1.6M0.18%+78513F-HRTimeline
CWM, LLCin group CWM, LLC72,213$1.6M0.01%+9,46813F-HRTimelineGroup
DEUTSCHE BANK AG\in group DEUTSCHE BANK AG\70,763$1.57M<0.01%-549,11713F-HRTimelineGroup
Old Port Advisors70,250$1.56M0.43%-1,79313F-HRTimeline
NISA INVESTMENT ADVISORS, LLC70,172$1.56M0.01%-9,21013F-HRTimeline
MUTUAL OF AMERICA CAPITAL MANAGEMENT LLCin group MUTUAL OF AMERICA CAPITAL MAN…69,722$1.55M0.02%-2,60213F-HRTimelineGroup
Oak Thistle LLC68,170$1.51M0.22%new13F-HRTimeline
Brevan Howard Capital Management LPin group Brevan Howard Capital Managem…66,701$1.48M0.02%new13F-HRTimelineGroup
AMALGAMATED BANK64,747$1.44M0.01%-46013F-HRTimeline
STRS OHIO64,165$1.42M0.01%+15,24813F-HRTimeline
JANUS HENDERSON GROUP PLCin group Janus Henderson62,690$1.39M<0.01%+113F-HRTimelineGroup
Amundiin group Amundi59,892$1.39M<0.01%-14,20013F-HRTimelineGroup
Federation des caisses Desjardins du Qu…61,259$1.36M0.01%+15,30013F-HRTimeline
HANCOCK WHITNEY CORP60,824$1.35M0.03%+17,47313F-HRTimeline
Police & Firemen's Retirement System of…59,909$1.33M0.01%+2,44013F-HRTimeline
STIFEL FINANCIAL CORPin group STIFEL FINANCIAL CORP59,848$1.33M<0.01%+56313F-HRTimelineGroup
Accordant Advisory Group Inc56,820$1.26M0.94%new13F-HRTimeline
WINNACLE WEALTH LLC56,820$1.26M0.94%new13F-HRTimeline
Advisors Asset Management, Inc.56,015$1.24M0.02%+3,66513F-HR/ATimeline
TEACHERS RETIREMENT SYSTEM OF THE STATE…51,579$1.14M0.01%013F-HRTimeline
Squarepoint Ops LLCin group Squarepoint Ops LLC51,564$1.14M<0.01%new13F-HR/ATimelineGroup
Corient Private Wealth LPgroup of 2 filers holding49,153$1.14M-4,015Timeline
PUBLIC EMPLOYEES RETIREMENT SYSTEM OF O…50,403$1.12M<0.01%+64313F-HRTimeline
TEACHER RETIREMENT SYSTEM OF TEXAS49,203$1.09M<0.01%-17,12413F-HRTimeline
YOUSIF CAPITAL MANAGEMENT, LLC47,407$1.05M0.01%-1,05013F-HRTimeline
Centiva Capital, LP44,289$982K0.06%-1,61113F-HRTimeline
Ensign Peak Advisors, Inc44,286$982K<0.01%013F-HRTimeline
ARIZONA STATE RETIREMENT SYSTEM43,266$960K0.01%+2,66313F-HRTimeline
WEALTH ENHANCEMENT ADVISORY SERVICES, L…40,364$932K<0.01%+3,89713F-HRTimeline
Comprehensive Money Management Services…40,126$890K0.68%+7,33113F-HRTimeline
Zurcher Kantonalbank (Zurich Cantonalba…39,703$881K<0.01%013F-HRTimeline
Louisiana State Employees Retirement Sy…39,390$874K0.02%-413F-HRTimeline
MQS Management LLC38,398$852K0.47%new13F-HRTimeline
Quantinno Capital Management LP37,745$837K<0.01%-13,71713F-HRTimeline
CIBC WORLD MARKETS CORP37,325$828K0.01%-1,93113F-HRTimeline
HORIZON KINETICS ASSET MANAGEMENT LLC37,146$824K0.01%-4,23613F-HRTimeline
SCHULHOFF & CO INC36,982$820K0.36%+1013F-HRTimeline
IQ EQ FUND MANAGEMENT (IRELAND) Ltd36,909$819K0.07%new13F-HRTimeline
Bollard Group LLC36,516$810K0.02%-59,11613F-HRTimeline
COMMONWEALTH OF PENNSYLVANIA PUBLIC SCH…36,021$799K<0.01%+2,43013F-HRTimeline
SEI INVESTMENTS COin group SEI INVESTMENTS CO35,067$778K<0.01%+33613F-HRTimelineGroup
LPL Financial LLC34,827$772K<0.01%+3,80913F-HRTimeline
EDENTREE ASSET MANAGEMENT Ltd34,757$770K0.21%+7,90813F-HRTimeline
Horizon Investments, LLC34,176$758K0.01%new13F-HRTimeline
STATE OF MICHIGAN RETIREMENT SYSTEM33,715$748K<0.01%+10013F-HRTimeline
ENVESTNET ASSET MANAGEMENT INCin group ENVESTNET ASSET MANAGEMENT INC33,333$739K<0.01%+10,55813F-HRTimelineGroup
Handelsbanken Fonder AB33,195$736K<0.01%-12,50013F-HRTimeline
Thrivent Financial for Lutheransin group Thrivent Financial for Luther…33,010$732K<0.01%+6313F-HR/ATimelineGroup
Graham Capital Management, L.P.32,318$717K0.02%-25,60813F-HRTimeline
TUDOR INVESTMENT CORP ET AL32,283$716K0.01%new13F-HRTimeline
NORTHWESTERN MUTUAL INVESTMENT MANAGEME…group of 2 filers holding32,086$712K+39Timeline
COMMONWEALTH EQUITY SERVICES, LLC31,891$707K<0.01%+4,72813F-HRTimeline
MAI Capital Management31,684$703K<0.01%+31,18313F-HRTimeline
NEW MEXICO EDUCATIONAL RETIREMENT BOARD31,632$702K0.02%013F-HRTimeline
KLP KAPITALFORVALTNING AS30,853$684K<0.01%+4,00013F-HRTimeline
LEAVELL INVESTMENT MANAGEMENT, INC.30,351$673K0.03%013F-HRTimeline
EXCHANGE TRADED CONCEPTS, LLC30,020$666K0.01%+57613F-HRTimeline
Mariner, LLCin group Mariner, LLC29,714$659K<0.01%+17,12213F-HRTimelineGroup
OREGON PUBLIC EMPLOYEES RETIREMENT FUND28,668$636K0.01%+10013F-HRTimeline
Summit Trail Advisors, LLC28,362$629K0.01%new13F-HRTimeline
BOK Financial Private Wealth, Inc.28,157$625K0.04%-1,14613F-HRTimeline
CARY STREET PARTNERS ASSET MANAGEMENT L…group of 2 filers holding27,880$618K+18,094Timeline
Creative Planningin group Creative Planning27,844$618K<0.01%+5,39813F-HRTimelineGroup
Foresight Group Ltd Liability Partnersh…27,696$613K0.09%013F-HRTimeline
CITIGROUP INCin group CITIGROUP INC27,540$611K<0.01%+49413F-HRTimelineGroup
Ameritas Investment Partners, Inc.group of 2 filers holding27,226$604K-423Timeline
Penn Mutual Asset Management27,000$599K1.06%+18,00013F-HRTimeline
Integrated Wealth Concepts LLC26,104$579K0.01%+8,13513F-HRTimeline
ExodusPoint Capital Management, LP25,132$557K0.01%new13F-HRTimeline
Elo Mutual Pension Insurance Co24,423$542K0.01%new13F-HRTimeline
PNC Financial Services Group, Inc.in group PNC Financial Services Group,…23,785$528K<0.01%-3,00913F-HRTimelineGroup
Painted Porch Advisors LLC23,750$527K0.22%013F-HRTimeline
BRIGHTON JONES LLC23,019$511K0.01%+14,97813F-HRTimeline
VOYA INVESTMENT MANAGEMENT LLCin group VOYA INVESTMENT MANAGEMENT LLC21,599$479K<0.01%+2,33313F-HRTimelineGroup
Pallas Capital Advisors LLC21,128$469K0.02%+3,26513F-HRTimeline
MONTGOMERY INVESTMENT MANAGEMENT INC20,544$456K0.18%+1613F-HRTimeline
New Age Alpha Advisors, LLC20,446$453K0.02%+46213F-HRTimeline
Mountain Hill Investment Partners Corp.20,440$453K0.38%+4,99313F-HRTimeline
Waverly Advisors, LLC20,315$451K<0.01%+3,05313F-HRTimeline
CAPITAL FUND MANAGEMENT S.A.20,193$448K<0.01%new13F-HRTimeline
XPONANCE LLC20,008$444K<0.01%-65413F-HRTimeline
Inspire Investing, LLCin group Inspire Investing, LLC19,854$440K0.05%+7,49713F-HRTimelineGroup
Groupe la Francaisein group Groupe la Francaise19,737$437K0.01%-79813F-HRTimelineGroup
Aster Capital Management (DIFC) Ltd19,567$434K0.1%+17,34813F-HRTimeline
DRW Securities, LLC18,846$418K0.03%new13F-HRTimeline
MERCER GLOBAL ADVISORS INC /ADV18,730$415K<0.01%+2,91913F-HRTimeline
INCEPTIONR LLC18,471$410K0.09%+3,15513F-HRTimeline
GABELLI FUNDS LLCin group GAMCO Investors (Gabelli)17,260$383K<0.01%013F-HRTimelineGroup
Clearstead Advisors, LLC17,194$381K0.01%-1,09613F-HRTimeline
LANDSCAPE CAPITAL MANAGEMENT, L.L.C.in group LANDSCAPE CAPITAL MANAGEMENT,…16,684$370K0.03%new13F-HRTimelineGroup
STATE OF WISCONSIN INVESTMENT BOARD16,600$368K<0.01%+2,06913F-HRTimeline
KEYBANK NATIONAL ASSOCIATION/OHin group KEYBANK NATIONAL ASSOCIATION/…16,510$366K<0.01%-1,17413F-HRTimelineGroup
NEUBERGER BERMAN GROUP LLCin group NEUBERGER BERMAN GROUP LLC16,349$363K<0.01%+2,20913F-HR/ATimelineGroup
PUBLIC EMPLOYEES RETIREMENT ASSOCIATION…15,785$350K<0.01%013F-HRTimeline
CAPTRUST FINANCIAL ADVISORS15,280$339K<0.01%-6,35613F-HRTimeline
Essential Partners LLC15,096$335K0.13%13F-HRTimeline
Versor Investments LP15,059$334K0.06%-50,45613F-HRTimeline
Timber Creek Capital Management LLC13,957$333K0.08%013F-HRTimeline
Nebula Research & Development LLC14,615$324K0.03%new13F-HRTimeline
OPPENHEIMER ASSET MANAGEMENT INC.in group OPPENHEIMER ASSET MANAGEMENT …14,390$319K<0.01%+3,38213F-HRTimelineGroup
CoreCommodity Management, LLC14,020$311K0.05%013F-HRTimeline
Kestra Investment Management, LLCin group Kestra Advisory Services, LLC13,846$307K0.01%+13,82413F-HR/ATimelineGroup
AXA S.A.in group AXA S.A.13,800$306K<0.01%+5,11513F-HRTimelineGroup
AlphaQuest LLC13,298$295K0.03%+8,43013F-HRTimeline
Hamilton Point Investment Advisors, LLC13,180$292K0.05%-313F-HRTimeline
Baker Ellis Asset Management LLC13,130$291K0.04%-20413F-HRTimeline
BLAIR WILLIAM & CO/IL13,113$291K<0.01%+4,24113F-HRTimeline
Dynamic Technology Lab Private Ltd12,605$280K0.04%new13F-HRTimeline
NATIXIS ADVISORS, LLCin group Natixis Investment Managers12,485$277K<0.01%new13F-HRTimelineGroup
TRUIST FINANCIAL CORPin group TRUIST FINANCIAL CORP12,071$268K<0.01%-2,91813F-HRTimelineGroup
SUSQUEHANNA INTERNATIONAL GROUP, LLPin group SUSQUEHANNA INTERNATIONAL GRO…11,971$266K<0.01%-13,25913F-HRTimelineGroup
BOOTHBAY FUND MANAGEMENT, LLC11,722$260K0.01%-255,64113F-HRTimeline
Abel Hall, LLC11,520$256K0.12%new13F-HRTimeline
State of Wyoming10,984$244K0.03%new13F-HRTimeline

Events

  • COHEN & STEERS, INC. and 4 other reporting persons reported 12,590,454 RYN shares, 4.23% of the class, as of 2026-09-30, in a 13G/A filed 2026-10-07. 0001284812-26-000188

    Accepted , ingested

  • Daniels Ryan M. sold 14 RYN shares on 2026-10-02, Form 4 filed 2026-10-06. 0002125295-26-000017

    Accepted , ingested