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Disclosery

PRGS PROGRESS SOFTWARE CORP

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Stock (common), issuer CIK 876167

301 managers (309 13F filers) reported a position in PRGS shares at 2025-09-30, 47,180,944 shares in all. 40 filers opened a position and 76 filers sold out during the quarter.

Managers
301
13F filers
309
New
40
Added
109
Reduced
120
Sold out
76
Shares held by filers
47,180,944

Shares held by filers adds up every 13F filer's report, so a share reported by a parent and by its subsidiary counts twice (H9). Managers counts each filer group once.

13D and 13G filers

No 13D or 13G on this security in the data. Structured 13D/G data starts 2024-12-18.

Insider trades

The latest Form 4 transactions, non-derivative.
OwnerDateSharesPriceFiling
WANG YUFAN STEPHANIEofficer (Chief Legal Officer)2026-10-05-1,245 S$37Form 4

13F holders, Q3 2025

Filers in the available 13F reports with a share position at 2025-09-30, filer groups collapsed to one row (method), largest first. Options and principal amounts are left out. A group row states no % of portfolio, since each member reports its own portfolio: the members' percents are listed under it. A value about 1,000 times or 1/1,000 of the price other filers imply is shown as filed but neither ranks the row nor enters a sum.

HolderSharesValue% of portfolioChangeSource
BlackRock, Inc.in group BlackRock, Inc.6,873,467$302M0.01%+104,42313F-HRTimelineGroup
VANGUARD GROUP INCgroup of 2 filers holding5,704,249$251M-115,591Timeline
Boston Trust Walden Corpin group Boston Trust Walden Corp1,950,676$85.7M0.63%+153,95713F-HRTimelineGroup
STATE STREET CORPin group STATE STREET CORP1,790,608$78.7M<0.01%-1,36313F-HRTimelineGroup
WELLINGTON MANAGEMENT GROUP LLPin group WELLINGTON MANAGEMENT GROUP L…1,348,134$59.2M0.01%+262,45413F-HRTimelineGroup
Allspring Global Investments Holdings, …in group Allspring Global Investments …1,178,716$53.3M0.09%+243,95013F-HRTimelineGroup
DIMENSIONAL FUND ADVISORS LPin group DIMENSIONAL FUND ADVISORS LP1,139,309$50M0.01%+20,66913F-HRTimelineGroup
GEODE CAPITAL MANAGEMENT, LLCin group GEODE CAPITAL MANAGEMENT, LLC1,075,392$47.2M<0.01%+7,93013F-HRTimelineGroup
SYSTEMATIC FINANCIAL MANAGEMENT LP914,798$40.2M0.98%+270,77313F-HRTimeline
MANUFACTURERS LIFE INSURANCE COMPANY, T…in group MANUFACTURERS LIFE INSURANCE …906,411$39.8M0.03%+230,46513F-HRTimelineGroup
Invesco Ltd.in group Invesco Ltd.868,245$38.1M0.01%+20,39713F-HRTimelineGroup
MACQUARIE GROUP LTDin group MACQUARIE GROUP LTD862,126$37.9M0.04%+24,10113F-HRTimelineGroup
PRINCIPAL FINANCIAL GROUP INCgroup of 2 filers holding843,739$37.1M-72,047Timeline
Swedbank ABin group Swedbank AB744,226$32.7M0.03%+21,50013F-HRTimelineGroup
FMR LLCin group FMR LLC734,265$32.3M<0.01%-45,96813F-HRTimelineGroup
Capital International Investorsin group Capital International Investo…722,081$31.7M0.01%013F-HRTimelineGroup
AMERICAN CENTURY COMPANIES INCin group AMERICAN CENTURY COMPANIES INC648,792$28.5M0.01%+550,14513F-HRTimelineGroup
SEGALL BRYANT & HAMILL, LLC643,145$28.3M0.38%+104,61113F-HRTimeline
FULLER & THALER ASSET MANAGEMENT, INC.638,215$28M0.1%-77513F-HRTimeline
BANK OF AMERICA CORP /DE/in group BANK OF AMERICA CORP /DE/630,137$27.7M<0.01%-74,21713F-HR/ATimelineGroup
BARCLAYS PLCin group BARCLAYS PLC514,999$22.6M0.01%-59,72313F-HR/ATimelineGroup
OAKTOP CAPITAL MANAGEMENT II, L.P.508,608$22.3M2.02%013F-HRTimeline
CHARLES SCHWAB INVESTMENT MANAGEMENT INCin group Charles Schwab Trust Co499,206$21.9M<0.01%-1,75613F-HRTimelineGroup
MORGAN STANLEYin group MORGAN STANLEY494,382$21.7M<0.01%-352,13213F-HR/ATimelineGroup
DekaBank Deutsche Girozentrale478,870$21M0.04%013F-HRTimeline
NORTHERN TRUST CORPin group NORTHERN TRUST CORP474,363$20.8M<0.01%-19,09113F-HRTimelineGroup
Diamond Hill Capital Management, LLC (I…464,598$20.4M0.1%+275,20513F-HRTimeline
DEUTSCHE BANK AG\in group DEUTSCHE BANK AG\431,499$19M0.01%-437,58413F-HRTimelineGroup
GOLDMAN SACHS GROUP INCin group GOLDMAN SACHS GROUP INC430,179$18.9M<0.01%-300,70313F-HRTimelineGroup
MILLENNIUM MANAGEMENT LLCin group MILLENNIUM MANAGEMENT LLC417,969$18.4M0.01%+7,32613F-HRTimelineGroup
Natixis Investment Managersgroup of 2 filers holding371,924$16.3M-184,280Timeline
Qube Research & Technologies Ltd371,593$16.3M0.02%new13F-HRTimeline
BANK OF MONTREAL /CAN/in group BANK OF MONTREAL /CAN/369,657$16.2M0.01%-232,77213F-HRTimelineGroup
LSV ASSET MANAGEMENT362,000$15.9M0.04%+234,10013F-HRTimeline
MARSHALL WACE, LLPin group MARSHALL WACE, LLP357,384$15.7M0.02%+314,45913F-HRTimelineGroup
RICE HALL JAMES & ASSOCIATES, LLC338,941$14.9M0.81%+202,30513F-HRTimeline
UBS Group AGgroup of 2 filers holding326,695$14.4M-151,717Timeline
Bank of New York Mellon Corpin group Bank of New York Mellon Corp322,909$14.2M<0.01%-24,03413F-HRTimelineGroup
Glenmede Investment Management, LP295,352$13M0.06%+50,52613F-HRTimeline
ARROWSTREET CAPITAL, LIMITED PARTNERSHIPin group ARROWSTREET CAPITAL, LIMITED …290,308$12.8M0.01%+5,37713F-HRTimelineGroup
MIZUHO FINANCIAL GROUP INCgroup of 2 filers holding290,219$12.7M-477,748Timeline
WELLS FARGO & COMPANY/MNin group WELLS FARGO & COMPANY/MN275,126$12.1M<0.01%-69,10313F-HRTimelineGroup
JPMORGAN CHASE & COin group JPMORGAN CHASE & CO270,954$11.9M<0.01%-149,47113F-HR/ATimelineGroup
NOMURA HOLDINGS INCin group NOMURA ASSET MANAGEMENT CO LTD269,938$11.9M0.05%+223,82813F-HRTimelineGroup
Swiss Life Asset Management Ltdin group Swiss Life Asset Management L…264,285$11.6M0.06%+78113F-HRTimelineGroup
KENNEDY CAPITAL MANAGEMENT LLC258,211$11.3M0.25%+82,07013F-HRTimeline
Heron Bay Capital Management254,311$11.2M2.09%+137,18913F-HRTimeline
Connor, Clark & Lunn Investment Managem…228,140$10M0.03%+3,60113F-HRTimeline
Jefferies Financial Group Inc.in group Jefferies Financial Group Inc.223,616$9.82M0.09%-337,50513F-HRTimelineGroup
AQR CAPITAL MANAGEMENT LLCin group AQR CAPITAL MANAGEMENT LLC214,738$9.43M0.01%+177,42213F-HRTimelineGroup
BNP PARIBAS FINANCIAL MARKETSin group BNP PARIBAS FINANCIAL MARKETS203,343$8.93M0.01%+160,70913F-HRTimelineGroup
Woodline Partners LP202,602$8.9M0.04%+41,28213F-HRTimeline
SUSQUEHANNA INTERNATIONAL GROUP, LLPgroup of 3 filers holding173,738$7.63M+28,917Timeline
Public Sector Pension Investment Board168,160$7.39M0.03%+1,91913F-HRTimeline
UniSuper Management Pty Ltd157,004$6.9M0.04%+14,78013F-HRTimeline
BANK OF NOVA SCOTIAin group BANK OF NOVA SCOTIA154,585$6.79M0.01%+8,77213F-HRTimelineGroup
PRUDENTIAL FINANCIAL INCin group PRUDENTIAL FINANCIAL INC149,181$6.55M0.01%+55,26513F-HRTimelineGroup
AMERIPRISE FINANCIAL INCin group AMERIPRISE FINANCIAL INC147,983$6.5M<0.01%-191,18913F-HRTimelineGroup
Edgestream Partners, L.P.142,466$6.26M0.18%+56,04013F-HRTimeline
TWO SIGMA INVESTMENTS, LPgroup of 2 filers holding135,526$5.95M+68,282Timeline
KLP KAPITALFORVALTNING AS127,100$5.58M0.02%+28,00013F-HRTimeline
RHUMBLINE ADVISERS125,082$5.49M<0.01%-1,07713F-HRTimeline
Universal- Beteiligungs- und Serviceges…in group Universal- Beteiligungs- und …124,552$5.47M0.01%-35,09813F-HRTimelineGroup
Nuveen, LLCin group TIAA123,391$5.42M<0.01%013F-HRTimelineGroup
Y-Intercept (Hong Kong) Ltd117,801$5.17M0.12%new13F-HRTimeline
Legal & General Group Plcin group Legal & General Group Plc115,413$5.07M<0.01%-6,68313F-HRTimelineGroup
JANE STREET GROUP, LLCin group JANE STREET GROUP, LLC105,592$4.64M0.01%-22,41113F-HRTimelineGroup
GHP Investment Advisors, Inc.95,251$4.18M0.18%-21,25013F-HRTimeline
Corient Private Wealth LPin group Corient Private Wealth LP94,149$4.14M0.01%-20,92413F-HRTimelineGroup
Hillsdale Investment Management Inc.90,950$4M0.12%+4,90013F-HRTimeline
FIRST TRUST ADVISORS LPin group FIRST TRUST ADVISORS LP84,712$3.72M<0.01%+32,67113F-HRTimelineGroup
Swiss National Bank83,600$3.67M<0.01%+1,40013F-HRTimeline
CITADEL ADVISORS LLCin group CITADEL ADVISORS LLC81,519$3.58M<0.01%-51,34113F-HRTimelineGroup
CAPITAL FUND MANAGEMENT S.A.77,822$3.42M0.03%+12,55113F-HRTimeline
RAYMOND JAMES FINANCIAL INCin group RAYMOND JAMES FINANCIAL INC75,307$3.31M<0.01%-53,97813F-HRTimelineGroup
ALLIANCEBERNSTEIN L.P.72,442$3.18M<0.01%-2,58913F-HRTimeline
CITIGROUP INCin group CITIGROUP INC70,425$3.09M<0.01%-246,69913F-HRTimelineGroup
STIFEL FINANCIAL CORPin group STIFEL FINANCIAL CORP69,994$3.07M<0.01%-98213F-HRTimelineGroup
DARK FOREST CAPITAL MANAGEMENT LP69,552$3.06M0.11%new13F-HR/ATimeline
GAMCO Investors (Gabelli)group of 2 filers holding69,408$3.05M+2,110Timeline
NEW YORK STATE TEACHERS RETIREMENT SYST…68,851$3.02M0.01%-5,33013F-HRTimeline
PRICE T ROWE ASSOCIATES INC /MD/65,936$2.9M<0.01%-160,72613F-HRTimeline
VICTORY CAPITAL MANAGEMENT INC62,282$2.74M<0.01%+17,55613F-HRTimeline
Choate Investment Advisors61,948$2.72M0.06%013F-HRTimeline
ENVESTNET ASSET MANAGEMENT INCgroup of 2 filers holding61,028$2.68M-38,092Timeline
GW&K Investment Management, LLC59,475$2.61M0.02%new13F-HRTimeline
RENAISSANCE TECHNOLOGIES LLC56,965$2.5M<0.01%-283,66413F-HRTimeline
ROYAL BANK OF CANADAin group ROYAL BANK OF CANADA54,982$2.42M<0.01%+9,62113F-HRTimelineGroup
F/m Investments LLC54,139$2.38M0.09%-26913F-HRTimeline
ROYCE & ASSOCIATES LP53,702$2.36M0.02%+11213F-HRTimeline
Campbell & CO Investment Adviser LLC52,919$2.32M0.18%+42,76913F-HRTimeline
ANCHOR CAPITAL ADVISORS LLC52,340$2.3M0.07%new13F-HRTimeline
CANADA LIFE ASSURANCE Coin group CANADA LIFE ASSURANCE Co48,215$2.06M<0.01%+60113F-HRTimelineGroup
Mariner, LLCin group Mariner, LLC45,834$2.01M<0.01%-10,16513F-HRTimelineGroup
PARADIGM CAPITAL MANAGEMENT INC/NY45,300$1.99M0.09%013F-HRTimeline
Van Hulzen Asset Management, LLC44,496$1.95M0.15%-49113F-HRTimeline
Polaris Capital Management, LLC44,200$1.94M0.13%+9,60013F-HRTimeline
MOUNTAIN PACIFIC INVESTMENT ADVISERS LLC42,658$1.87M0.29%-184,96813F-HRTimeline
Trexquant Investment LP42,166$1.85M0.02%new13F-HRTimeline
Russell Investments Group, Ltd.in group Russell Investments Group, Lt…41,810$1.84M<0.01%+4,74313F-HRTimelineGroup
FRANKLIN RESOURCES INCin group Franklin Resources41,487$1.82M<0.01%+63913F-HRTimelineGroup
SEI INVESTMENTS COin group SEI INVESTMENTS CO40,428$1.78M<0.01%-12,90313F-HRTimelineGroup
CALIFORNIA STATE TEACHERS RETIREMENT SY…39,641$1.74M<0.01%+213F-HRTimeline
AXA Investment Managers S.A.37,141$1.63M<0.01%13F-HRTimeline
QSV Equity Investors LLC35,883$1.58M1.61%013F-HRTimeline
Brevan Howard Capital Management LPin group Brevan Howard Capital Managem…33,202$1.46M0.02%new13F-HRTimelineGroup
Gotham Asset Management, LLC32,692$1.44M0.01%+27,46413F-HRTimeline
PEAK6 LLCin group PEAK6 LLC32,056$1.41M0.04%+31,79913F-HR/ATimelineGroup
Verition Fund Management LLC31,655$1.39M0.01%+11,54913F-HRTimeline
Thrivent Financial for Lutheransin group Thrivent Financial for Luther…31,099$1.37M<0.01%-15,89213F-HRTimelineGroup
TEACHER RETIREMENT SYSTEM OF TEXAS30,527$1.34M0.01%+11,56613F-HRTimeline
Daiwa Securities Group Inc.in group Daiwa Securities Group Inc.30,000$1.32M<0.01%+23,70013F-HRTimelineGroup
MACKENZIE FINANCIAL CORP29,939$1.32M<0.01%+14,84013F-HRTimeline
Arax Advisory Partners28,697$1.26M0.02%+28,67513F-HRTimeline
BROWN ADVISORY INCin group BROWN ADVISORY INC28,680$1.26M<0.01%-10,00013F-HRTimelineGroup
MOODY NATIONAL BANK TRUST DIVISIONin group MOODY NATIONAL BANK TRUST DIV…27,873$1.22M0.1%+5,45713F-HRTimelineGroup
HUDSON EDGE INVESTMENT PARTNERS INC.27,214$1.2M0.19%+10,52313F-HRTimeline
D. E. Shaw & Co., Inc.in group D. E. Shaw & Co., Inc.25,892$1.14M<0.01%-160,73913F-HRTimelineGroup
HSBC HOLDINGS PLCin group HSBC HOLDINGS PLC25,797$1.13M<0.01%-73,44313F-HRTimelineGroup
Squarepoint Ops LLCin group Squarepoint Ops LLC25,629$1.13M<0.01%-48,65413F-HRTimelineGroup
MetLife Investment Management, LLCin group MetLife Investment Management…25,353$1.11M0.01%013F-HRTimelineGroup
Caption Management, LLC25,200$1.11M0.06%-40,45313F-HRTimeline
INTECH INVESTMENT MANAGEMENT LLC25,165$1.11M0.01%-19,68813F-HRTimeline
Elo Mutual Pension Insurance Co24,828$1.09M0.02%+9,63413F-HRTimeline
Illinois Municipal Retirement Fund24,539$1.08M0.01%+1,81213F-HRTimeline
SG Americas Securities, LLC23,649$1.04M<0.01%+14,87613F-HRTimeline
VIRGINIA RETIREMENT SYSTEMS ET Al22,400$984K0.01%+17,90013F-HRTimeline
State of Alaska, Department of Revenue21,691$952K0.01%-3,93713F-HRTimeline
DELPHI MANAGEMENT INC /MA/21,296$936K0.87%+5913F-HRTimeline
Integrated Investment Consultants, LLC21,279$935K0.16%+13,02213F-HRTimeline
OPPENHEIMER & CO INCin group OPPENHEIMER ASSET MANAGEMENT …20,348$894K0.01%new13F-HRTimelineGroup
MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.in group Mirae Asset Global Investment…19,906$874K<0.01%-3,57913F-HRTimelineGroup
Louisiana State Employees Retirement Sy…19,800$870K0.01%-30013F-HRTimeline
CastleKnight Management LP19,700$865K0.05%+12,80013F-HRTimeline
JANUS HENDERSON GROUP PLCin group Janus Henderson19,257$845K<0.01%013F-HRTimelineGroup
Holocene Advisors, LP18,831$827K<0.01%new13F-HRTimeline
EXCHANGE TRADED CONCEPTS, LLC18,533$814K0.01%-15,35513F-HRTimeline
Zurcher Kantonalbank (Zurich Cantonalba…17,521$770K<0.01%+6,55713F-HRTimeline
Creative Planningin group Creative Planning17,318$761K<0.01%-2,93613F-HRTimelineGroup
Pathstone Holdings, LLCin group Pathstone Holdings, LLC16,216$712K<0.01%+3,67013F-HRTimelineGroup
ADVISORY RESEARCH INC15,209$668K0.09%+56213F-HRTimeline
Cerity Partners LLCin group Cerity Partners LLC14,816$651K<0.01%+6,64313F-HRTimelineGroup
Walleye Trading LLCin group Walleye Capital LLC14,518$638K0.01%-11,53913F-HRTimelineGroup
COMERICA BANK14,268$627K<0.01%-15,58713F-HRTimeline
NEW YORK STATE COMMON RETIREMENT FUND13,972$614K<0.01%013F-HRTimeline
Kendall Capital Management13,930$612K0.14%-5,14013F-HRTimeline
PIER 88 INVESTMENT PARTNERS LLC13,670$601K0.11%new13F-HRTimeline
YOUSIF CAPITAL MANAGEMENT, LLC13,339$586K0.01%-2,75113F-HRTimeline
HARBOR CAPITAL ADVISORS, INC.13,272$583K0.05%+1,51313F-HRTimeline
ARIZONA STATE RETIREMENT SYSTEM12,743$560K<0.01%+19213F-HRTimeline
AMALGAMATED BANK12,671$557K<0.01%+42213F-HRTimeline
STATE BOARD OF ADMINISTRATION OF FLORID…12,376$544K<0.01%013F-HRTimeline
TEACHERS RETIREMENT SYSTEM OF THE STATE…12,269$539K<0.01%-96013F-HRTimeline
WEDBUSH SECURITIES INC12,094$531K0.02%+7,47013F-HRTimeline
Inspire Investing, LLCgroup of 2 filers holding12,056$530K+1,459Timeline
Police & Firemen's Retirement System of…11,858$521K<0.01%+2,10013F-HRTimeline
Bayesian Capital Management, LP11,606$510K0.1%+7,48713F-HRTimeline
Handelsbanken Fonder AB11,516$506K<0.01%013F-HRTimeline
Tidal Investments LLCin group Tidal Investments LLC11,142$489K<0.01%-1,25713F-HRTimelineGroup
COMMONWEALTH OF PENNSYLVANIA PUBLIC SCH…10,978$482K<0.01%-6813F-HRTimeline
Engineers Gate Manager LP10,832$476K0.01%new13F-HRTimeline
INCEPTIONR LLC10,715$471K0.11%new13F-HRTimeline
State of Tennessee, Department of Treas…10,658$468K<0.01%013F-HRTimeline
CAPSTONE INVESTMENT ADVISORS, LLCin group CAPSTONE INVESTMENT ADVISORS,…10,638$467K<0.01%new13F-HRTimelineGroup
Ensign Peak Advisors, Inc10,044$441K<0.01%-7,95013F-HR/ATimeline
CHOREO, LLC10,032$441K0.01%-5,18113F-HRTimeline
Grantham, Mayo, Van Otterloo & Co. LLC10,016$440K<0.01%-7,11213F-HRTimeline
PNC Financial Services Group, Inc.in group PNC Financial Services Group,…9,984$439K<0.01%+1,69113F-HRTimelineGroup
OREGON PUBLIC EMPLOYEES RETIREMENT FUND9,672$425K0.01%-20013F-HRTimeline
Cetera Investment Advisers9,525$418K<0.01%+5,24213F-HRTimeline
Blueshift Asset Management, LLC9,503$417K0.13%new13F-HRTimeline
XTX Topco Ltdin group XTX Topco Ltd9,502$417K0.02%+1,71213F-HRTimelineGroup
Quantinno Capital Management LP9,330$410K<0.01%+2,83713F-HRTimeline
QRG CAPITAL MANAGEMENT, INC.8,711$383K<0.01%-3,46113F-HRTimeline
OXFORD ASSET MANAGEMENT LLP8,467$372K0.12%new13F-HRTimelineThe filing omitted confidential holdings, a later amendment may add them.
Amundiin group Amundi7,964$372K<0.01%-147,56213F-HRTimelineGroup
Freestone Grove Partners LP8,279$364K<0.01%-2,94413F-HRTimeline
ProShare Advisors LLC8,075$355K<0.01%-1,49213F-HRTimeline
JAMES INVESTMENT RESEARCH, INC.7,818$343K0.04%+1,84513F-HRTimeline
VOYA INVESTMENT MANAGEMENT LLCin group VOYA INVESTMENT MANAGEMENT LLC7,725$339K<0.01%-4,13813F-HRTimelineGroup
DANSKE BANK A/S7,633$335K<0.01%-12,71713F-HR/ATimeline
BOOTHBAY FUND MANAGEMENT, LLC7,531$331K0.01%new13F-HRTimeline
SIMPLEX TRADING, LLC7,477$328K0.01%-43,24113F-HRTimeline
Graham Capital Management, L.P.7,438$327K0.01%new13F-HRTimeline
FORBES J M & CO LLP7,120$313K0.03%013F-HRTimeline
Atria Investments, Inc7,062$310K<0.01%+1,45413F-HRTimeline
Ifrah Financial Services, Inc.6,925$304K0.08%+9613F-HRTimeline
AXQ CAPITAL, LP6,747$296K0.11%new13F-HRTimeline
AlphaQuest LLC6,568$289K0.02%+22513F-HRTimeline
Virtu Financial LLCin group Virtu Financial LLC6,442$283K0.02%+3,15213F-HRTimelineGroup
PRAXIS INVESTMENT MANAGEMENT INC6,410$282K0.02%013F-HRTimeline
Maryland State Retirement & Pension Sys…6,269$275K0.01%-6,38013F-HRTimeline
LAZARD ASSET MANAGEMENT LLCin group LAZARD ASSET MANAGEMENT LLC6,152$270K<0.01%-32,47213F-HRTimelineGroup
Versor Investments LP6,100$268K0.05%-3,60013F-HRTimeline
Orion Porfolio Solutions, LLC6,051$266K<0.01%+73613F-HRTimeline
Pictet Asset Management Holding SAin group Pictet Asset Management Holdi…5,960$262K<0.01%013F-HRTimelineGroup
DRW Securities, LLC5,697$251K0.01%new13F-HRTimeline
HighTower Advisors, LLC5,631$247K<0.01%-3,19413F-HRTimeline
CenterBook Partners LP5,612$247K0.01%new13F-HRTimeline
Henshaw Capital LLC5,528$243K0.01%-2,02913F-HRTimeline

Events

  • WANG YUFAN STEPHANIE sold 1,245 PRGS shares on 2026-10-05, Form 4 filed 2026-10-07. 0000876167-26-000127

    Accepted , ingested