GREAT LAKES ADVISORS, LLC: IONS position timeline

GREAT LAKES ADVISORS, LLC held 66,924 IONS shares at 2026-06-30 per its 13F-HR filed 2026-08-10.

the latest 13F is the newest step

1. period ended2026-06-30

   13F 66,924 shares

   Change vs previous 13F -52,002 shares

   0.04% of the 13F portfolio, value $5,306,404.

   13F-HR 0001078013-26-000004 filed by GREAT LAKES ADVISORS, LLC
   Filed 2026-08-10 (acceptance time not in the bulk data set), ingested 2026-10-06 09:57:27 UTC

   Acceptance time not in the bulk data set (spec §4.2).

2. period ended2026-03-31

   13F 118,926 shares

   Change vs previous 13F -30,254 shares

   0.07% of the 13F portfolio, value $8,930,153.

   13F-HR 0001078013-26-000003 filed by GREAT LAKES ADVISORS, LLC
   Filed 2026-05-12 (acceptance time not in the bulk data set), ingested 2026-10-06 09:58:43 UTC

   Acceptance time not in the bulk data set (spec §4.2).

3. period ended2025-12-31

   13F 149,180 shares

   Change vs previous 13F -37,770 shares

   0.1% of the 13F portfolio, value $11,801,630.

   13F-HR 0001078013-26-000002 filed by GREAT LAKES ADVISORS, LLC
   Filed 2026-02-12 (acceptance time not in the bulk data set), ingested 2026-10-07 03:01:36 UTC

   Acceptance time not in the bulk data set (spec §4.2).

4. period ended2025-09-30

   13F 186,950 shares

   Change vs previous 13F -212,696 shares

   0.1% of the 13F portfolio, value $12,230,269.

   13F-HR 0001078013-25-000008 filed by GREAT LAKES ADVISORS, LLC
   Filed 2025-11-03 (acceptance time not in the bulk data set), ingested 2026-10-07 03:02:25 UTC

   Acceptance time not in the bulk data set (spec §4.2).

5. period ended2025-06-30

   13F 399,646 shares

   Change vs previous 13F not available

   0.13% of the 13F portfolio, value $15,790,013.

   13F-HR 0001078013-25-000005 filed by GREAT LAKES ADVISORS, LLC
   Filed 2025-08-11 (acceptance time not in the bulk data set), ingested 2026-10-07 18:23:11 UTC

   Acceptance time not in the bulk data set (spec §4.2).