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13F-HR filed by ProShare Advisors LLC

ProShare Advisors LLC filed this 13F-HR for the quarter ended 2021-03-31, filed 2021-05-14, with 2,257 holding rows worth $31,802,827,000.

Accession
0001085146-21-001681
Filed
2021-05-14
Accepted by EDGAR
acceptance time not in the bulk data set
Ingested
Source
SEC quarterly data set

Original filing

13F report for Q1 2021

A holdings report, 2,257 entries on the cover page, a total value of $31,802,827,000 stated on the cover page (in dollars after the unit check, H1), rows summing to $31,802,827,000 with VALUE read as thousands by filing date, 13F file number 028-12122. The quarter's positions are built on this report. The resolved quarter.

The information table after summing rows of one CUSIP and put or call (H4), with the CUSIP and issuer name as filed. Values in dollars after the unit check (method).

Issuer as filedCUSIPClassAmountValueValue read asType
KINIKSA PHARMACEUTICALS LTDG5269C101COM CL A11,560$214,000thousands by filing date
RE MAX HLDGS INC75524W108CL A5,420$213,000thousands by filing date
OPTIMIZERX CORP68401U204COM NEW4,361$213,000thousands by filing date
KOPPERS HOLDINGS INC50060P106COM6,125$213,000thousands by filing date
CITI TRENDS INC17306X102COMMON STOCK2,538$213,000thousands by filing date
LUMBER LIQUIDATORS HOLDINGS55003T107COMMON STOCK8,486$213,000thousands by filing date
USA TODAY CO INC36472T109COM39,463$212,000thousands by filing date
HORIZON BANCORP IND440407104COM11,427$212,000thousands by filing date
BRIGHTCOVE INC10921T101COM10,543$212,000thousands by filing date
NOVOCURE LTDG6674U108ORD SHS1,597$211,000thousands by filing date
FRANCHISE GROUP INC35180X105COM5,833$211,000thousands by filing date
ARGAN INC04010E109COM3,939$210,000thousands by filing date
MYERS INDUSTRIES INC628464109COMMON STOCK10,635$210,000thousands by filing date
1ST SOURCE CORP COM336901103Stock4,385$209,000thousands by filing date
IVERIC BIO INC46583P102COM33,861$209,000thousands by filing date
QAD INC-A74727D306CL A3,136$209,000thousands by filing date
ACCO BRANDS CORP COM00081T108Stock24,684$208,000thousands by filing date
TRUSTCO BK CORP N Y898349105COM28,161$208,000thousands by filing date
UNIQURE NVN90064101SHS6,157$207,000thousands by filing date
TUTOR PERINI CORP901109108COM10,919$207,000thousands by filing date
VIRTU FINL INC928254101CL A6,667$207,000thousands by filing date
SPARTANNASH CO847215100COM10,558$207,000thousands by filing date
CIRCOR INTL INC17273K109COM5,924$206,000thousands by filing date
SPECTRUM PHARMACEUTICALS INC84763A108COM63,046$206,000thousands by filing date
IDEAYA BIOSCIENCES INC COM45166A102Stock8,740$205,000thousands by filing date
COTY INC222070203COM CL A22,724$205,000thousands by filing date
WORLD WRESTLING ENTMT INC98156Q108CL A3,776$205,000thousands by filing date
ANAPTYSBIO INC032724106COM9,507$205,000thousands by filing date
EAGLE PHARMACEUTICALS INC269796108COM4,918$205,000thousands by filing date
HEIDRICK & STRUGGLES INTL IN422819102COM5,698$204,000thousands by filing date
PRECISION BIOSCIENCES INC74019P108COM19,685$204,000thousands by filing date
HUTCHMED CHINA LTD SPONSORED ADS44842L103ADR7,189$203,000thousands by filing date
CORSAIR GAMING INC22041X102COM6,108$203,000thousands by filing date
GENESCO INC371532102COM4,284$203,000thousands by filing date
SFL CORP LTDG7738W106COMMON STOCK25,308$203,000thousands by filing date
LIBERTY GLOBAL PLCG5480U104SHS CL A7,922$203,000thousands by filing date
AXOGEN INC05463X106COM9,974$202,000thousands by filing date
ENERGY FUELS INC292671708COM NEW35,615$202,000thousands by filing date
NICOLET BANKSHARES INC65406E102COM2,419$202,000thousands by filing date
AKEBIA THERAPEUTICS INC00972D105COMMON STOCK59,537$202,000thousands by filing date
CAI INTERNATIONAL INC12477X106COM4,447$202,000thousands by filing date
SP PLUS CORP78469C103COM6,151$202,000thousands by filing date
ELEMENT SOLUTIONS INC COM28618M106Stock10,942$200,000thousands by filing date
FIRST BANCSHARES INC MISS318916103COM5,452$200,000thousands by filing date
ANTERO MIDSTREAM CORP COM03676B102Stock22,013$199,000thousands by filing date
MODINE MFG CO COM607828100Stock13,396$198,000thousands by filing date
ENERGY RECOVERY INC29270J100COM10,721$197,000thousands by filing date
SPORTSMANS WHSE HLDGS INC84920Y106COM11,437$197,000thousands by filing date
MARINUS PHARMACEUTICALS INC56854Q200COM NEW12,716$197,000thousands by filing date
CYTOMX THERAPEUTICS INC23284F105COMMON STOCK25,408$196,000thousands by filing date
GRANITE PT MTG TR INC38741L107COM STK16,222$194,000thousands by filing date
TECTONIC THERAPEUTIC INC COM05455M100Stock15,278$194,000thousands by filing date
THERAPEUTICSMD INC88338N107COM143,790$193,000thousands by filing date
COLONY CR REAL ESTATE INC19625T101COM CL A22,467$191,000thousands by filing date
INSEEGO CORP45782B104COMMON STOCK19,079$191,000thousands by filing date
MRC GLOBAL INC55345K103COM21,000$190,000thousands by filing date
Heritage Commerce Corp426927109Common Stock15,541$190,000thousands by filing date
WIDEOPENWEST INC96758W101COM13,983$190,000thousands by filing date
GERON CORP374163103COM119,736$189,000thousands by filing date
HARBORONE BANCORP INC NEW41165Y100COM NEW13,886$187,000thousands by filing date
MURPHY OIL CORP COM626717102Stock11,195$184,000thousands by filing date
BRIGHTVIEW HLDGS INC10948C107COM10,933$184,000thousands by filing date
ATRECA INC04965G109COM11,863$182,000thousands by filing date
CHATHAM LODGING TR16208T102COM13,775$181,000thousands by filing date
ANTARES PHARMA INC036642106COM43,922$181,000thousands by filing date
TPG RE FIN TR INC COM87266M107REIT16,065$180,000thousands by filing date
CRINETICS PHARMACEUTICALS INC COM22663K107Stock11,786$180,000thousands by filing date
AMNEAL PHARMACEUTICALS INC03168L105COM STK CL A26,524$179,000thousands by filing date
MEIRAGTX HOLDINGS PLCG59665102COMMON STOCK12,382$179,000thousands by filing date
DHT HOLDINGS INCY2065G121SHS NEW29,795$177,000thousands by filing date
CROSSFIRST BANKSHARES INC22766M109COM12,807$177,000thousands by filing date
CAPSTEAD MTG CORP14067E506COM NO PAR28,333$177,000thousands by filing date
NEOLEUKIN THERAPEUTICS INC64049K104COM14,411$177,000thousands by filing date
ELLINGTON FINANCIAL INC28852N109COM11,022$176,000thousands by filing date
SUNCOKE ENERGY INC86722A103COMMON STOCK24,635$173,000thousands by filing date
FOSSIL GROUP INC34988V106COMMON STOCK13,918$173,000thousands by filing date
VERASTEM INC92337C104COM69,083$171,000thousands by filing date
ECOVYST INC.73943T103Common Stock10,257$171,000thousands by filing date
PAR PAC HOLDINGS INC69888T207COM NEW11,872$168,000thousands by filing date
RAYONIER ADVANCED MATLS INC75508B104COM18,425$167,000thousands by filing date
FRANKLIN STR PPTYS CORP35471R106COM30,564$167,000thousands by filing date
BRIGHAM MINERALS INC10918L103COMMON STOCK11,335$166,000thousands by filing date
POSEIDA THERAPEUTICS INC73730P108COM17,434$166,000thousands by filing date
FLEXION THERAPEUTICS INC33938J106COM18,504$166,000thousands by filing date
ENTERCOM COMMUNICATIONS CORP293639100CL A31,342$165,000thousands by filing date
BOINGO WIRELESS INC09739C102COM11,706$165,000thousands by filing date
CORNERSTONE BLDG BRANDS INC21925D109COM11,712$164,000thousands by filing date
LIBERTY LATIN AMERICA LTDG9001E102COM CL A12,607$162,000thousands by filing date
MITEK SYS INC606710200COM NEW11,105$162,000thousands by filing date
PG&E CORP COM69331C108Stock13,780$161,000thousands by filing date
NEXTIER OILFIELD SOLUTIONS INC65290C105COMMON STOCK43,241$161,000thousands by filing date
SIGA TECHNOLOGIES INC COM826917106Stock24,639$160,000thousands by filing date
NEOPHOTONICS CORP64051T100COM13,265$159,000thousands by filing date
CONSOLIDATED COMMUNICATIONS209034107COM21,978$158,000thousands by filing date
SYROS PHARMACEUTICALS INC87184Q107COM21,119$158,000thousands by filing date
Veru Inc92536C103Common Stock14,611$157,000thousands by filing date
AKOUOS INC00973J101COM11,286$157,000thousands by filing date
RIBBON COMMUNICATIONS INC762544104COMMON STOCK18,837$155,000thousands by filing date
GLOBAL MED REIT INC37954A204COM NEW11,747$154,000thousands by filing date
A10 NETWORKS INC002121101COM16,010$154,000thousands by filing date
METALLUS INC887399103COMMON STOCK13,135$154,000thousands by filing date
ASPIRA WOMENS HEALTH INC04537Y109COM22,759$154,000thousands by filing date
ORBCOMM INC68555P100COM20,198$154,000thousands by filing date
CHIMERIX INC COM16934W106Stock15,901$153,000thousands by filing date
URANIUM ENERGY CORP COM916896103Stock52,495$150,000thousands by filing date
VBI VACCINES INC91822J103COM NEW48,155$150,000thousands by filing date
SELECTA BIOSCIENCES INC816212104COM33,068$150,000thousands by filing date
ARLO TECHNOLOGIES INC04206A101COMMON STOCK23,541$148,000thousands by filing date
AEGLEA BIOTHERAPEUTICS INC00773J103COM18,697$148,000thousands by filing date
BIODELIVERY SCIENCES INTL IN09060J106COM37,969$148,000thousands by filing date
TENNECO INC880349105COM13,683$147,000thousands by filing date
ACCEL ENTERTAINMENT INC00436Q106COMMON STOCK13,491$147,000thousands by filing date
EXPRO GROUP HOLDINGS NV COMN33462107Stock41,411$147,000thousands by filing date
PACTIV EVERGREEN INC69526K105COM10,570$145,000thousands by filing date
DIANTHUS THERAPEUTICS INC COM55910K108Stock12,214$145,000thousands by filing date
TRANSPORTATION & LOGISTI438083107COM15,356$144,000thousands by filing date
GOGO INC38046C109COM14,743$142,000thousands by filing date
PAE INC69290Y109COM CL A15,785$142,000thousands by filing date
DORIAN LPG LTDY2106R110SHS USD10,661$140,000thousands by filing date
Viewra, Inc92672L107COM32,288$140,000thousands by filing date
ORCHID IS CAP INC68571X103COM22,888$138,000thousands by filing date
ATHENEX INC04685N103COM31,895$137,000thousands by filing date
NEW SR INVT GROUP INC648691103COM21,902$136,000thousands by filing date
TIDEWATER INC NEW COM88642R109Stock10,800$135,000thousands by filing date
PUMA BIOTECHNOLOGY INC74587V107COMMON STOCK13,765$134,000thousands by filing date
CATCHMARK TIMBER TR INC CL A14912Y202REIT13,026$133,000thousands by filing date
CROSS CTRY HEALTHCARE INC227483104COM10,598$132,000thousands by filing date
TRUECAR INC89785L107COM27,499$132,000thousands by filing date
COSTAMARE INCY1771G102SHS13,509$130,000thousands by filing date
ONESPAWORLD HOLDINGS LIMITEDP73684113COM12,076$129,000thousands by filing date
KALA BIO INC COM483119103Stock19,079$129,000thousands by filing date
NORDIC AMERICAN TANKERS LIMIG65773106COM39,328$128,000thousands by filing date
MBIA INC55262C100COMMON STOCK13,267$128,000thousands by filing date
APARTMENT INVT & MGMT CO CL A03748R747REIT20,747$127,000thousands by filing date
CYMABAY THERAPEUTICS INC23257D103COM28,039$127,000thousands by filing date
PARATEK PHARMACEUTICALS INC699374302Equity17,971$127,000thousands by filing date
PREFERRED APT CMNTYS INC74039L103COM12,714$125,000thousands by filing date
LIBERTY TRIPADVISOR HLDGS IN531465102COM SER A19,424$124,000thousands by filing date
AXT INC00246W103COM10,549$123,000thousands by filing date
ACCURAY INC DEL004397105COM24,551$122,000thousands by filing date
CITY OFFICE REIT INC178587101COM11,440$121,000thousands by filing date
CATALYST PHARMACEUTICALS INC14888U101COM25,933$120,000thousands by filing date
CONTANGO OIL & GAS CO COM NEW21075N204Stock30,727$120,000thousands by filing date
DURECT CORP266605104COM59,592$118,000thousands by filing date
CHICOS FAS INC168615102COM35,302$117,000thousands by filing date
WHITESTONE REIT966084204COM11,828$115,000thousands by filing date
VYNE THERAP.92941V209COM16,787$115,000thousands by filing date
EDGIO INC53261M104COM32,176$115,000thousands by filing date
AGENUS INC00847G705COM NEW42,022$114,000thousands by filing date
CALAMP CORP128126109COMMON STOCK10,357$112,000thousands by filing date
JOUNCE THERAPEUTICS INC481116101COM10,872$112,000thousands by filing date
EIGER BIOPHARMACEUTICALS INC28249U105COM12,674$112,000thousands by filing date
VIKING THERAPEUTICS INC92686J106COM17,524$111,000thousands by filing date
ZIX CORP98974P100COM14,601$110,000thousands by filing date
MONEYGRAM INTL INC60935Y208COM NEW16,596$109,000thousands by filing date
OIL STS INTL INC678026105COM17,866$108,000thousands by filing date
FREQUENCY THERAPEUTICS INC35803L108COM11,394$108,000thousands by filing date
HERSHA HOSPITALITY TR427825500PR SHS BEN INT10,176$107,000thousands by filing date
TELLURIAN INC NEW87968A104COM44,919$105,000thousands by filing date
GREENSKY INC39572G100CL A16,878$104,000thousands by filing date
NIAGEN BIOSCIENCE INC171077407COM NEW11,047$103,000thousands by filing date
DASEKE INC23753F107COM12,174$103,000thousands by filing date
MEI PHARMA INC55279B202COM NEW29,306$101,000thousands by filing date
SIENTRA INC82621J105COM13,844$101,000thousands by filing date
UNITY BIOTECHNOLOGY INC91381U101COM16,776$101,000thousands by filing date
Berry Corp08579X101Common Stock18,094$100,000thousands by filing date
ADAPTIMMUNE THERAPEUTICS PLC00653A107SPONDS ADR18,840$100,000thousands by filing date
COMPUGEN LTDM25722105ORD11,576$99,000thousands by filing date
ONCOCYTE CORP68235C107COM19,048$99,000thousands by filing date
CYTOSORBENTS CORP23283X206CMN10,975$95,000thousands by filing date
COREPOINT LODGING INC21872L104COM10,493$95,000thousands by filing date
VIRNETX HLDG CORP92823T108COM16,946$94,000thousands by filing date
QUANTUM CORP747906501COM11,113$93,000thousands by filing date
RPC INC749660106COMMON STOCK17,078$92,000thousands by filing date
MEDICINOVA INC58468P206COM NEW18,203$92,000thousands by filing date
XERIS PHARMACEUTICALS INC98422L107COM20,464$92,000thousands by filing date
W & T OFFSHORE INC92922P106COM25,194$90,000thousands by filing date
STANDARD BIOTOOLS INC34385P108COM19,768$89,000thousands by filing date
KEZAR LIFE SCIENCES INC49372L100COM14,766$88,000thousands by filing date
IBIO INC451033203COM NEW57,196$88,000thousands by filing date
WAVE LIFE SCIENCES LTD SHSY95308105Stock15,536$87,000thousands by filing date
OVID THERAPEUTICS INC690469101COMMON STOCK21,489$86,000thousands by filing date
ACACIA RESEARCH CORP003881307COMMON STOCK12,714$85,000thousands by filing date
VAXART INC NEW92243A200COM14,001$85,000thousands by filing date
ATHERSYS INC04744L106COM46,345$83,000thousands by filing date
INVACARE CORP461203101COMMON STOCK10,040$81,000thousands by filing date
STEREOTAXIS INC85916J409COMMON STOCK11,810$79,000thousands by filing date
NN INC629337106COM11,204$79,000thousands by filing date
SELECT WATER SOLUTIONS INC81617J301CL A COM15,747$78,000thousands by filing date
LEXICON PHARMACEUTICALS INC528872302COMMON STOCK13,289$78,000thousands by filing date
EROS INTL PLCG3788M114SHS NEW43,359$78,000thousands by filing date
GTY TECHNOLOGY HOLDINGS INC362409104COM12,278$78,000thousands by filing date
CITIZENS INC174740100COMMON STOCK13,247$77,000thousands by filing date
NATIONAL CINEMEDIA INC635309107COM16,636$77,000thousands by filing date
TRICIDA INC89610F101COM14,506$77,000thousands by filing date
NPK INTERNATIONAL INC651718504COMMON STOCK23,992$75,000thousands by filing date
PAVMED INC70387R106COM16,683$74,000thousands by filing date
RADIANT LOGISTICS INC75025X100COM10,452$73,000thousands by filing date
EZCORP INC302301106CL A NON VTG14,474$72,000thousands by filing date
NEWAGE INC650194103COMMON STOCK25,346$72,000thousands by filing date
QUOTIENT LTDG73268107SHS19,366$71,000thousands by filing date
DAKTRONICS INC234264109COM10,872$68,000thousands by filing date
EXPRESS INC30219E103COMMON STOCK16,990$68,000thousands by filing date
ITERIS INC NEW46564T107COM11,040$68,000thousands by filing date
SUMMIT THERAPEUTICS INC86627T108COM11,425$67,000thousands by filing date
ATLANTIC PWR CORP04878Q863COM NEW23,243$67,000thousands by filing date
CHIASMA INC16706W102COM21,385$67,000thousands by filing date
CALITHERA BIOSCIENCES INC COM NEW13089P101Stock27,402$66,000thousands by filing date
WAITR HLDGS INC930752100COM22,366$66,000thousands by filing date
ENTRAVISION COMMUNICATIONS A29382R107COMMON STOCK15,717$63,000thousands by filing date
FORTRESS BIOTECH INC34960Q109COM17,861$63,000thousands by filing date
CONCERT PHARMACEUTICALS INC206022105COM12,344$62,000thousands by filing date
ASSEMBLY BIOSCIENCES INC045396108COM13,472$62,000thousands by filing date
COMSCORE INC20564W105COM16,183$59,000thousands by filing date
APPLIED GENETIC TECHNOL CORP03820J100COM11,430$58,000thousands by filing date
RESONANT INC76118L102COM13,628$58,000thousands by filing date
SURGALIGN HOLDINGS INC86882C105COM26,527$58,000thousands by filing date
LIVEXLIVE MEDIA INC53839L208COM NEW13,344$58,000thousands by filing date
VOYAGER THERAPEUTICS INC92915B106COMMON STOCK12,116$57,000thousands by filing date
HARVARD BIOSCIENCE INC416906105COM10,229$56,000thousands by filing date
PEABODY ENGR CORP704551100COM16,929$52,000thousands by filing date
HC2 HLDGS INC404139107COM13,223$52,000thousands by filing date
WESTERN ASSET MTG CAP CORP95790D105COM15,907$51,000thousands by filing date
GOLD RESOURCE CORP38068T105COMMON STOCK17,912$47,000thousands by filing date
Fluent Co34380C102COM11,175$46,000thousands by filing date
REPUBLIC FIRST BANCORP INC760416107COM12,146$46,000thousands by filing date
ACELRX PHARMACEUTICALS INC00444T100COM26,826$46,000thousands by filing date
ALASKA COMMUNICATIONS SYS GR01167P101COM14,003$46,000thousands by filing date
FALCON MINERALS CORP30607B109CL A COM10,203$46,000thousands by filing date
MUSTANG BIO INC62818Q104COM13,455$45,000thousands by filing date
MILESTONE PHARMACEUTICALS IN59935P209COM NEW12,297$44,000thousands by filing date
DHI GROUP INC23331S100COM12,824$43,000thousands by filing date
Checkpoint Therapeutics Inc Com162828107COM13,829$43,000thousands by filing date
CASI PHARMACEUTICALS INC14757U109COM17,666$42,000thousands by filing date
Enzo Biochem Inc294100102Common Stock11,869$41,000thousands by filing date
PIXELWORKS INC72581M305COM NEW12,175$40,000thousands by filing date
CORBUS PHARMACEUTICALS HLDGS21833P103COM20,523$40,000thousands by filing date
Arbutus Biopharma Corp03879J100Common Shares, without par value11,750$39,000thousands by filing date
RVL PHARMACEUTICALS PLCG6S41R101COM12,053$39,000thousands by filing date
MARRONE BIO INNOVATIONS INC57165B106COM18,530$39,000thousands by filing date
SYNCHRONOSS TECHNOLOGIES INC87157B103COM10,548$38,000thousands by filing date
AGILE THERAPEUTICS INC00847L100COM18,146$38,000thousands by filing date
OVERSEAS SHIPHOLDING GROUP I69036R863CL A NEW17,640$36,000thousands by filing date
PIERIS PHARMACEUTICALS INC720795103COM14,179$36,000thousands by filing date
EVOFEM BIOSCIENCES INC30048L104COM20,490$36,000thousands by filing date
SERVICESOURCE INTL LLC81763U100COM23,573$35,000thousands by filing date
EXICURE INC30205M101COM15,848$35,000thousands by filing date
CERECOR INC15671L109COM11,495$35,000thousands by filing date
SAFE BULKERS INCY7388L103COMMON STOCK13,866$34,000thousands by filing date
TYME TECHNOLOGIES INC COM90238J103Stock18,562$33,000thousands by filing date
STRONGBRIDGE BIOPHARMA PLCG85347105SHS USD11,763$32,000thousands by filing date
ADMA BIOLOGICS INC000899104COM17,612$31,000thousands by filing date
ABEONA THERAPEUTICS INC00289Y107COM16,134$30,000thousands by filing date
SAVARA INC805111101COM12,833$27,000thousands by filing date
NYMOX PHARMACEUTICAL CORP ORD SHS CorpP73398102Common Stock11,299$25,000thousands by filing date
ROCKWELL MED INC774374102COM18,622$22,000thousands by filing date
SOLENO THERAPEUTIVS, INC.834203200COMM16,124$20,000thousands by filing date

Amendment chain

FilingRoleFiledAcceptedIngested
13F-HR 0001085146-21-001681this filing2021-05-14acceptance time not in the bulk data set