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13F-HR filed by TWO SIGMA INVESTMENTS, LP

TWO SIGMA INVESTMENTS, LP filed this 13F-HR for the quarter ended 2021-03-31, filed 2021-05-17, with 3,749 holding rows worth $38,436,761,000.

Accession
0000919574-21-003689
Filed
2021-05-17
Accepted by EDGAR
acceptance time not in the bulk data set
Ingested
Source
SEC quarterly data set

Original filing

13F report for Q1 2021

A holdings report, 3,749 entries on the cover page, a total value of $38,436,761,000 stated on the cover page (in dollars after the unit check, H1), rows summing to $38,436,761,000 with VALUE read as thousands by filing date, 13F file number 028-10385. The quarter's positions are built on this report. The resolved quarter.

Other managers

The information table after summing rows of one CUSIP and put or call (H4), with the CUSIP and issuer name as filed. Values in dollars after the unit check (method).

Issuer as filedCUSIPClassAmountValueValue read asType
USA TRUCK INC902925106COM12,694$243,000thousands by filing date
PAGERDUTY INC COM69553P100Stock6,000$241,000thousands by filing dateput
ZIMMER BIOMET HOLDINGS INC COM98956P102Stock1,500$240,000thousands by filing dateput
TECK RESOURCES LTD CL B878742204Stock12,500$240,000thousands by filing datecall
OLD DOMINION FREIGHT LINE INC COM679580100Stock1,000$240,000thousands by filing dateput
American Tower Corporation03027X100COM1,000$239,000thousands by filing dateput
American Tower Corporation03027X100COM1,000$239,000thousands by filing datecall
ADT INC DEL COM00090Q103Stock28,305$239,000thousands by filing date
BURLINGTON STORES INC COM122017106Stock800$239,000thousands by filing datecall
GUARANTY BANCSHARES INC400764106COMMON STOCK6,507$239,000thousands by filing date
DEUTSCHE BK AG NAMEN AKTD18190898Stock19,808$238,000thousands by filing date
FRESHPET INC COM358039105Stock1,500$238,000thousands by filing datecall
SURO CAP CORP86887Q109COM NEW17,558$238,000thousands by filing date
CONNECTONE BANCORP INC20786W107COMMON STOCK9,390$238,000thousands by filing date
HILTON GRAND VACATIONS INC COM43283X105Stock6,300$236,000thousands by filing dateput
BANK OF NY MELLON CORP COM064058100Stock5,000$236,000thousands by filing datecall
UNITED NAT FOODS INC COM911163103Stock7,163$236,000thousands by filing date
MACERICH CO554382101COM20,200$236,000thousands by filing datecall
DANA INC235825205COM9,701$236,000thousands by filing date
STATE AUTO FINANCIAL CORP855707105COM11,959$236,000thousands by filing date
SERVICESOURCE INTL LLC81763U100COM160,174$235,000thousands by filing date
IDT CORP448947507CL B NEW10,334$234,000thousands by filing date
FIRST BANCSHARES INC MISS318916103COM6,400$234,000thousands by filing date
XCELENERGY INC98389B100COM3,500$233,000thousands by filing datecall
DELEK US HLDGS INC NEW COM24665A103Stock10,715$233,000thousands by filing date
REPLIGEN CORP COM759916109Stock1,200$233,000thousands by filing dateput
BYLINE BANCORP INC124411109COM11,013$233,000thousands by filing date
COMPASS DIVERSIFIED HOLDINGS20451Q104COMMON STOCK10,062$233,000thousands by filing date
CONNS INC208242107COM12,000$233,000thousands by filing datecall
LANDOS BIOPHARMA INC515069102CMN24,152$233,000thousands by filing date
BUILDERS FIRSTSOURCE INC COM12008R107Stock5,000$232,000thousands by filing dateput
NORTHERN TR CORP COM665859104Stock2,200$231,000thousands by filing datecall
STMICROELECTRONICS N V NY REGISTRY861012102ADR6,026$231,000thousands by filing date
COCA COLA CONS INC191098102COM801$231,000thousands by filing date
RING ENERGY INC76680V108COMMON STOCK100,101$231,000thousands by filing date
SUNOPTA INC8676EP108COM15,628$231,000thousands by filing date
TD SYNNEX CORPORATION COM87162W100Stock2,000$230,000thousands by filing datecall
MICROVISION INC DEL594960304COM NEW12,400$230,000thousands by filing datecall
SUMMIT MATERIALS INC CLASS A86614U100CL A8,200$230,000thousands by filing dateput
CONTINENTAL RES INC212015101COM8,900$230,000thousands by filing dateput
NATURAL GAS SERVICES GROUP63886Q109COMMON STOCK24,386$230,000thousands by filing date
GROUPON INC399473206COM NEW4,531$229,000thousands by filing date
PEOPLES BANCORP INC709789101COM6,900$229,000thousands by filing date
CALIX INC13100M509COM6,600$229,000thousands by filing date
ATRICURE INC04963C209COM3,500$229,000thousands by filing date
PARK OHIO HOLDINGS CORP700666100COMMON STOCK7,266$229,000thousands by filing date
RETRACTABLE TECHNOLOGIES INC76129W105COM17,900$229,000thousands by filing datecall
HOME PT CAPITAL INCORPORATED43734L106COM24,596$229,000thousands by filing date
KINGSOFT CLOUD HLDGS LTD ADS49639K101ADR5,800$228,000thousands by filing dateput
NATIONAL BK HLDGS CORP633707104CL A5,745$228,000thousands by filing date
CHURCHILL DOWNS INC171484108COM1,000$227,000thousands by filing datecall
BERRY GLOBAL GROUP INC08579W103COM3,700$227,000thousands by filing datecall
CURO GROUP HOLDINGS CORP23131L107COM15,570$227,000thousands by filing date
MOTOROLA SOLUTIONS INC COM NEW620076307Stock1,200$226,000thousands by filing datecall
BRIGHTHOUSE FINL INC10922N103COM5,100$226,000thousands by filing datecall
Up Fintech Holding Ltd91531W106COM12,700$226,000thousands by filing dateput
FOOT LOCKER INC344849104COM4,000$225,000thousands by filing dateput
OPTIMUM COMMUNICATIONS INC02156K103CL A6,900$224,000thousands by filing dateput
EXTRA SPACE STORAGE INC COM30225T102REIT1,683$223,000thousands by filing date
MATTEL INC577081102COM11,200$223,000thousands by filing dateput
NANOSTRING TECHNOLOGIES INC63009R109COM3,400$223,000thousands by filing dateput
ILLINOIS TOOL WKS INC452308109COM1,000$222,000thousands by filing datecall
PRINCIPAL FINANCIAL GROUP INC COM74251V102Stock3,700$222,000thousands by filing dateput
NOVAGOLD RESOURCES INC66987E206COM NEW25,300$222,000thousands by filing dateput
STEWART INFORMATION SVCS COR860372101COM4,276$222,000thousands by filing date
DARLING INGREDIENTS INC COM237266101Stock3,000$221,000thousands by filing datecall
VISTRA CORP COM92840M102Stock12,500$221,000thousands by filing dateput
AEROVIRONMENT INC COM008073108Stock1,900$221,000thousands by filing dateput
AMN HEALTHCARE SVCS INC001744101COM3,000$221,000thousands by filing datecall
VERITEX HLDGS INC923451108COM6,761$221,000thousands by filing date
AGNC INVT CORP COM00123Q104REIT13,100$220,000thousands by filing dateput
ABERCROMBIE & FITCH CO CL A002896207Stock6,400$220,000thousands by filing datecall
JACK IN THE BOX INC466367109COM2,000$220,000thousands by filing dateput
ANTERO MIDSTREAM CORP COM03676B102Stock24,200$219,000thousands by filing datecall
LUMENTUM HLDGS INC COM55024U109Stock2,400$219,000thousands by filing datecall
AGENUS INC00847G705COM NEW80,524$219,000thousands by filing date
Charles Schwab Corp808513105COM3,349$218,000thousands by filing date
CULLEN FROST BANKERS INC COM229899109Stock2,000$218,000thousands by filing dateput
CULLEN FROST BANKERS INC COM229899109Stock2,000$218,000thousands by filing datecall
Honeywell Technologies438516106COM1,000$217,000thousands by filing dateput
EQUIFAX INC COM294429105Stock1,200$217,000thousands by filing dateput
DAVITA INC COM23918K108Stock2,000$216,000thousands by filing datecall
WESTWOOD HOLDINGS GROUP INC961765104COMMON STOCK14,951$216,000thousands by filing date
EXPEDITORS INTL WASH INC COM302130109Stock2,000$215,000thousands by filing datecall
HYATT HOTELS CORP COM CL A448579102Stock2,600$215,000thousands by filing datecall
YUM CHINA HLDGS INC98850P109COM3,638$215,000thousands by filing date
EVENTBRITE INC29975E109COM CL A9,700$215,000thousands by filing datecall
ECOLAB INC COM278865100Stock1,000$214,000thousands by filing dateput
ECOLAB INC COM278865100Stock1,000$214,000thousands by filing datecall
ANNALY CAPITAL MANAGEMENT035710409COM24,900$214,000thousands by filing dateput
AXCELLA HEALTH INC05454B105COM44,901$214,000thousands by filing date
ROCKWELL AUTOMATION INC COM773903109Stock801$213,000thousands by filing date
WESTAMERICA BANCORPORATION COM957090103Stock3,398$213,000thousands by filing date
PAPA JOHNS INTL INC698813102COM2,400$213,000thousands by filing dateput
ZOGENIX INC.98978L204Common Stock10,917$213,000thousands by filing date
PAYCOMSOFTWAREINC70432V102STOK574$212,000thousands by filing date
TORM PLC SHS CL AG89479102Stock23,300$212,000thousands by filing date
DELUXE CORP MEDIUM TERM NTS COM248019101Stock5,064$212,000thousands by filing date
GUESS INC COM401617105Stock9,000$212,000thousands by filing dateput
XP INC CL AG98239109Stock5,600$211,000thousands by filing dateput
BARRETT BUSINESS SVCS INC068463108COM3,060$211,000thousands by filing date
CITRIX SYSTEMS INC177376100COM1,500$211,000thousands by filing dateput
LIVEPERSON, INC.538146101COM4,000$211,000thousands by filing datecall
HIMAX TECHNOLOGIES INC SPONSORED ADR43289P106ADR15,400$210,000thousands by filing datecall
UNITED STATES STL CORP NEW912909108COM8,000$209,000thousands by filing datecall
WEX INC96208T104COM1,000$209,000thousands by filing dateput
Fluent Co34380C102COM51,000$209,000thousands by filing date
ASSURED GUARANTY LTD COMG0585R106Stock4,927$208,000thousands by filing date
GLOBANT S AL44385109COM1,000$208,000thousands by filing datecall
VTI922908769COM1,000$207,000thousands by filing dateput
VTI922908769COM1,000$207,000thousands by filing datecall
SELECTQUOTE INC816307300COM7,000$207,000thousands by filing date
RUHNN HLDG LTD781314109ADS61,891$207,000thousands by filing date
NOVAGOLD RESOURCES INC66987E206COM NEW23,500$206,000thousands by filing datecall
HANGER INC41043F208COMMON STOCK9,044$206,000thousands by filing date
CALITHERA BIOSCIENCES INC COM NEW13089P101Stock85,000$206,000thousands by filing date
HAWAIIAN HOLDINGS INC419879101COM7,665$204,000thousands by filing date
L3HARRIS TECHNOLOGIES INC COM502431109Stock1,000$203,000thousands by filing datecall
DELEK US HLDGS INC NEW COM24665A103Stock9,300$203,000thousands by filing dateput
KEURIG DR PEPPER INC COM49271V100Stock5,898$203,000thousands by filing date
LADDER CAP CORP CL A505743104REIT17,209$203,000thousands by filing date
SANDERSON FARMS800013104COM1,300$203,000thousands by filing datecall
GRACELL BIOTECHNOLOGIES INC38406L103ADR13,174$203,000thousands by filing date
FISERV INC337738108COM1,695$202,000thousands by filing date
NEW YORK TIMES CO MTN BE CL A650111107Stock4,000$202,000thousands by filing datecall
LENNAR CORP CL A526057104Stock2,000$202,000thousands by filing datecall
GRAHAM CORP384556106COM14,180$202,000thousands by filing date
NXP SEMICONDUCTORS N V COMN6596X109Stock1,000$201,000thousands by filing datecall
FLUOR CORP343412102COM8,700$201,000thousands by filing datecall
TREAN INS GROUP INC89457R101COM12,467$201,000thousands by filing date
ACADIA RLTY TR004239109COM SH BEN INT10,524$200,000thousands by filing date
CONTANGO OIL & GAS CO COM NEW21075N204Stock51,171$200,000thousands by filing date
VERRICA PHARMACEUTICALS INC92511W108COM13,100$198,000thousands by filing date
SIGA TECHNOLOGIES INC COM826917106Stock30,227$196,000thousands by filing date
LIBERTY ENERGY INC53115L104COM CL A17,229$195,000thousands by filing date
HUYA INC ADS EACH REP 1 ORD SHS CL A44852D108COM10,000$195,000thousands by filing dateput
LUMEN TECHNOLOGIES INC550241103COM14,500$194,000thousands by filing datecall
MACERICH CO554382101COM16,500$193,000thousands by filing dateput
GREAT ELM CAP CORP390320109COM56,685$193,000thousands by filing date
Galiano Gold Inc36352H100COM171,593$192,000thousands by filing date
CARTER BANKSHARES INC146103106COMMON STOCK13,500$188,000thousands by filing date
OPKO HEALTH INC68375N103COM43,704$187,000thousands by filing date
CONTAINER STORE GROUP INC210751103COMMON STOCK11,100$185,000thousands by filing dateput
FREQUENCY THERAPEUTICS INC35803L108COM19,500$185,000thousands by filing date
NEOLEUKIN THERAPEUTICS INC64049K104COM14,887$183,000thousands by filing date
INFINERA CORP45667G103COMMON STOCK18,800$181,000thousands by filing datecall
RLJ LODGING TR COM74965L101REIT11,589$179,000thousands by filing date
LUCIRA HEALTH INC54948U105COM14,816$179,000thousands by filing date
BLUEROCK RESIDENTIAL GWT REI09627J102COM CL A17,725$179,000thousands by filing date
INSEEGO CORP45782B104COMMON STOCK17,395$174,000thousands by filing date
CBDMD INC12482W101COM41,769$173,000thousands by filing date
COTY INC222070203COM CL A19,100$172,000thousands by filing datecall
CALYXT INC13173L107COM28,431$171,000thousands by filing date
ACCO BRANDS CORP COM00081T108Stock20,039$169,000thousands by filing date
RIBBON COMMUNICATIONS INC762544104COMMON STOCK20,283$167,000thousands by filing date
GALERA THERAPEUTICS EQUICLASS EQUITY36338D108COMMON STOCK18,894$167,000thousands by filing date
EXICURE INC30205M101COM76,132$166,000thousands by filing date
MBIA INC55262C100COMMON STOCK17,152$165,000thousands by filing date
FULCRUM THERAPEUTICS INC359616109COMMON STOCK13,979$165,000thousands by filing date
STRONGBRIDGE BIOPHARMA PLCG85347105SHS USD59,400$164,000thousands by filing date
LSI INDUSTRIES INC50216C108COMMON STOCK19,084$163,000thousands by filing date
FULL HSE RESORTS INC359678109COM19,209$163,000thousands by filing date
TENNECO INC880349105COM15,100$162,000thousands by filing datecall
ARBOR REALTY TRUST INC038923108COM10,100$161,000thousands by filing dateput
ACACIA RESEARCH CORP003881307COMMON STOCK24,015$160,000thousands by filing date
APPLIED OPTOELECTRONICS INC03823U102COM18,600$155,000thousands by filing dateput
GOLAR LNG LTDG9456A100SHS15,200$155,000thousands by filing dateput
GASLOG PARTNERS LPY2687W108UNIT LTD PTNRP57,652$154,000thousands by filing date
LUMEN TECHNOLOGIES INC550241103COM11,300$151,000thousands by filing dateput
CLOUDERA INC18914U100COM12,400$151,000thousands by filing dateput
WHEELER REAL ESTATE INVT TR963025705COM NEW39,980$150,000thousands by filing date
PRIMIS FINANCIAL CORP74167B109COMMON STOCK10,272$149,000thousands by filing date
TRINITY BIOTECH PLC896438306SPON ADR NEW39,316$148,000thousands by filing date
GOPRO INC38268T103CL A12,600$147,000thousands by filing dateput
Enzo Biochem Inc294100102Common Stock42,859$147,000thousands by filing date
AEGLEA BIOTHERAPEUTICS INC00773J103COM18,332$145,000thousands by filing date
INFINERA CORP45667G103COMMON STOCK14,800$143,000thousands by filing dateput
ORCHARD THERAPEUTICS PLC68570P101ADR19,683$143,000thousands by filing date
BLUE APRON HLDGS INC09523Q200COM22,600$143,000thousands by filing dateput
ENDEAVOUR SILVER CORP29258Y103COM28,701$142,000thousands by filing date
WESTWATER RES INC961684206COM NEW26,000$140,000thousands by filing datecall
FIVE STAR SENIOR LIVING INC33832D205COM NEW22,792$139,000thousands by filing date
ALPHA PRO TECH LTD020772109COM14,200$139,000thousands by filing datecall
POLYPID LTDM8001Q118SHS15,039$138,000thousands by filing date
SURO CAP CORP86887Q109COM NEW10,000$136,000thousands by filing dateput
ORION ENERGY SYSTEMS INC686275108COM19,500$136,000thousands by filing date
XINYUAN REAL ESTATE CO LTD98417P105SPONS ADR48,433$136,000thousands by filing date
GEOSPACE TECHNOLOGIES CORP37364X109COM14,832$135,000thousands by filing date
ATHERSYS INC04744L106COM75,000$135,000thousands by filing date
VILLAGE FARMS INTL INC92707Y108COM10,000$132,000thousands by filing datecall
KINROSS GOLD CORP496902404COM19,500$130,000thousands by filing datecall
Zedge Inc98923T104COM10,700$129,000thousands by filing date
SILVERBOW RES INC82836G102COM16,222$126,000thousands by filing date
LUMOS PHARMA INC55028X109COM10,519$123,000thousands by filing date
PALATIN TECHNOLOGIES INC696077403COM PAR $ .01174,652$120,000thousands by filing date
GOLAR LNG LTDG9456A100SHS11,311$116,000thousands by filing date
OPERA LTD SPONSORED ADS68373M107ADR11,422$114,000thousands by filing date
MILLENDO THERAPEUTICS INC60040X103COM94,722$114,000thousands by filing date
BUILD-A-BEAR WORKSHOP INC120076104COM16,199$112,000thousands by filing date
ARC DOCUMENT SOLUTIONS INC00191G103COM53,246$112,000thousands by filing date
KANDI TECHNOLOGIES GROUP INC483709101COM16,850$105,000thousands by filing date
GRUPO FINANCIERO GALICIA399909100SPONSORED ADR13,600$103,000thousands by filing datecall
ObsEva SAH5861P103COM31,400$102,000thousands by filing date
Yatra Online Incorporated Ord SHSG98338109Common Stock42,375$99,000thousands by filing date
LIQTECH INTL INC53632A201COM12,119$98,000thousands by filing date
EQUITRANS MIDSTREAM CORP NPV294600101COM11,900$97,000thousands by filing dateput
EURODRY LTDY23508107COM10,984$96,000thousands by filing date
NOKIA CORP654902204SPONSORED ADR24,012$95,000thousands by filing date
HELIX ENERGY SOLUTIONS GRP INC COM42330P107Stock18,856$95,000thousands by filing date
WAVE LIFE SCIENCES LTD SHSY95308105Stock16,600$93,000thousands by filing date
LSB INDS INC502160104COM17,673$91,000thousands by filing date
PIERIS PHARMACEUTICALS INC720795103COM35,429$91,000thousands by filing date
CYCLACEL PHARMACE 6percent PFDCONV23254L603Stock12,855$91,000thousands by filing date
LONGEVERON INC CL A NEW54303L104Stock13,100$86,000thousands by filing date
ENTASIS THERAPEUTICS HLDGS I293614103COM39,361$84,000thousands by filing date
PHX MINERALS INC CL A69291A100Stock29,000$83,000thousands by filing date
BRICKELL BIOTECH INC10802T105COM73,285$80,000thousands by filing date
ACELRX PHARMACEUTICALS INC00444T100COM46,180$79,000thousands by filing date
BLUE APRON HLDGS INC09523Q200COM12,300$78,000thousands by filing datecall
LIFEWAY FOODS INC531914109COMMON STOCK14,285$77,000thousands by filing date
ARLO TECHNOLOGIES INC04206A101COMMON STOCK12,200$77,000thousands by filing dateput
LESAKA TECHNOLOGIES INC64107N206CMN13,228$74,000thousands by filing date
YPF SOCIEDAD ANONIMA984245100SPON ADR CL D17,778$73,000thousands by filing date
AMERISERV FINL INC03074A102COM18,070$73,000thousands by filing date
WRAP TECHNOLOGIES INC98212N107COM13,200$73,000thousands by filing dateput
ELEVATE CREDIT INC28621V101COM24,925$73,000thousands by filing date
WESTWATER RES INC961684206COM NEW13,500$72,000thousands by filing dateput
SYNACOR INC871561106COM32,362$71,000thousands by filing date
PERSPECTIVE THERAPEUTICS INC46489V104COM63,494$69,000thousands by filing date
SOUTHWESTERN ENERGY CO845467109COM14,000$65,000thousands by filing datecall
CHINA CUST RELATIONS CNTR ING2118P102COM10,744$65,000thousands by filing date
AILERON THERAPEUTICS INC COM NEW00887A105Stock45,100$64,000thousands by filing date
WASHINGTON PRIME GROUP NEW93964W405COM26,968$60,000thousands by filing date
CONSUMER PORTFOLIO SERVICES210502100COMMON STOCK14,708$59,000thousands by filing date
WRAP TECHNOLOGIES INC98212N107COM10,500$58,000thousands by filing datecall
GALMED PHARMACEUTICALS LTDM47238106SHS16,700$57,000thousands by filing date
CHIASMA INC16706W102COM18,147$57,000thousands by filing date
Allena Pharmaceuticals, Inc.018119107COM40,200$56,000thousands by filing date
OPKO HEALTH INC68375N103COM12,100$52,000thousands by filing dateput
APPLIED GENETIC TECHNOL CORP03820J100COM10,286$52,000thousands by filing date
SENSUS HEALTHCARE INC81728J109COM12,652$48,000thousands by filing date
OPKO HEALTH INC68375N103COM10,800$46,000thousands by filing datecall
SOCIETAL CDMO INC75629F109COM15,000$42,000thousands by filing date
FUEL TECH INC359523107COM12,388$39,000thousands by filing date
ELECTROCORE INC28531P103COM18,284$38,000thousands by filing date
IDEANOMICS INC45166V106COM11,322$33,000thousands by filing date
DIRTT ENVIRONMENTAL SOLUT LT25490H106COM10,612$33,000thousands by filing date
TITAN MED INC88830X819COM18,745$32,000thousands by filing date
GRAN TIERRA ENERGY INC38500T101COM32,224$23,000thousands by filing date

Amendment chain

FilingRoleFiledAcceptedIngested
13F-HR 0000919574-21-003689this filing2021-05-17acceptance time not in the bulk data set